The commercefacades extension exposes a group of facades which simplify the communication between the store frontend and the service layer. This is a summary which I wrote while studying for the certification. The Hybris Documentation about commercefacades has one page and it is worth reading.
The interaction of the controllers, facades, services, DAOs, models and data objects is explained in the this diagram: Simple Request Interaction Diagram.jpg
In the B2C Accelerator the converters and the populators are never used to convert from a DTO to a model. But they could be used.
–Based on Hybris 6.3